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doi:10.1016/j.neuron.2006.06.016   &n= bsp;3D"How  =20

Minireview

Drosophila Neuroblast Asymmetric Cell Division: Recent = Advances and=20 Implications for Stem Cell Biology

Fengwei Yu1,=20 2,=20 Chay T. Kuo1=20 and Yuh Nung Jan1,=20 ,
1Howard = Hughes=20 Medical Institute and Departments of Physiology, Biochemistry, and = Biophysics,=20 University of California, Rock Hall, 1550 Fourth Street, San Francisco,=20 California 94143
2Temasek Life Sciences = Laboratory and Department of Biological Sciences, National University of = Singapore, 1 Research Link, 117604, Singapore

Available online 5 = July=20 2006.

Asymmetric cell division is an evolutionarily conserved mechanism = widely used=20 to generate cellular diversity during development. Drosophila = neuroblasts=20 have been a useful model system for studying the molecular mechanisms of = asymmetric cell division. In this minireview, we focus on recent = progress in=20 understanding the role of heterotrimeric G proteins and their regulators = in=20 asymmetric spindle geometry, as well as the role of an = Inscuteable-independent=20 microtubule pathway in asymmetric localization of proteins in = neuroblasts. We=20 also discuss issues of progenitor proliferation and differentiation = associated=20 with asymmetric cell division and their broader implications for stem = cell=20 biology.


Article Outline

Main=20 Text
Receptor-Independent=20 Activation of Heterotrimeric G Proteins in Drosophila NBs
Heterotrimeric=20 G Proteins and Asymmetric Spindle Geometry
Asymmetric=20 Protein Localization and Mitotic Spindle Orientation
Asymmetric=20 Cell Division Coordinates Proliferation and Differentiation
Implications=20 for Mammalian Stem Cell Biology
References

Display=20 Full Size version of this image = (38K)

Figure 1. Asymmetric Cell Division of Drosophila NB

A schematic representation of NB delaminating from epithelium. As the = NB is=20 delaminating from the epithelium, it forms an apical stalk, which is in = contact=20 with the surrounding epithelial cells. During prophase, = Par/Insc/Pins/G=CE=B1i=20 proteins (red crescent) are asymmetrically localized at the apical = cortex,=20 whereas Numb/Pon/Pros/Mira (yellow crescent) proteins are localized at = the basal=20 cortex. The two centrosomes and the mitotic spindle (green) are aligned = parallel=20 to the surface of epithelium. During metaphase, the mitotic spindle = rotates 90=C2=B0=20 to become aligned perpendicular to the epithelial layer. During=20 anaphase/telophase, asymmetric spindle geometry is generated, which = divides the=20 neuroblast into two daughter cells of distinct fate and size: a large = apical NB=20 and a smaller basal GMC. The basal daughter GMC preferentially inherits=20 cell-fate determinants Numb and Pros.
Display=20 Full Size version of this image = (33K)

Figure 2. Canonical and Noncanonical Models of Heterotrimeric G = Protein=20 Signaling

(A) In the canonical model, upon ligand binding, a seven-pass = transmembrane=20 protein, GPCR, can act as a GEF to activate heterotrimeric G proteins. = In the=20 Drosophila NBs, heterotrimeric G proteins can also be activated = in a=20 receptor-independent manner.

(B) In a possible noncanonical cycle, two GDIs, Pins, Loco, and a = putative=20 cytosolic GEF, Ric-8, may act synergistically to activate heterotrimeric = G=20 proteins. The attenuation of G proteins may be mediated by Loco acting = as a GAP.=20
Display=20 Full Size version of this image = (45K)

Figure 3. Both Insc/Par and Dlg/Khc-73 Are Required for = Pins/G=CE=B1i=20 Crescent Formation and Spindle Alignment

In the wild-type NB, Insc/Par (red crescent) and Pins/G=CE=B1i (blue = crescent) are=20 colocalized at the apical cortex during prophase/metaphase and overlie = the=20 apical centrosome. In insc mutant NBs, during prophase = Pins/G=CE=B1i (blue) is=20 localized uniformly around the cortex and becomes a randomly placed = crescent=20 overlying one centrosome during metaphase. In dlg or = khc-73-RNAi=20 mutants, the Pins/G=CE=B1i crescent forms during prophase/metaphase, but = the mitotic=20 spindle becomes uncoupled during metaphase. In mutant NBs lacking=20 insc/dlg or insc/khc-73, Pins/G=CE=B1i fails = to form a=20 crescent during metaphase, suggesting that Pins crescent formation and = spindle=20 alignment require both pathways. Centrosomes are represented as green = dots.
Display=20 Full Size version of this image = (46K)

Figure 4. Asymmetrically Segregated Proteins aPKC and Brat/Pros = Can=20 Regulate Proliferation/Differentiation of Daughter Cells Following = Larval NB=20 Division

In the wild-type situation (top row), during cell division a larval = NB=20 segregates aPKC (red) to a proliferating NB daughter cell, and Brat/Pros = (yellow) to a differentiating GMC daughter cell. Brat/Pros can suppress = the=20 expression of cell cycle genes and dMyc in the GMC daughter cell, = thereby=20 inducing differentiation. In the double mutants of lgl/pins or=20 overexpression of a membrane-tethered aPKC (middle row), aPKC is = segregated into=20 both daughter cells, leading to a transformation of GMC into = proliferating NB.=20 In the brat or pros mutant larval NBs (bottom row), cell = cycle=20 progression and protein synthesis are not inhibited in the GMC, also = leading to=20 the transformation of GMC into proliferating NB.


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POSITION: absolute; TOP: -11px; HEIGHT: 14px; = BACKGROUND-COLOR: #777777 } .libBannerText { FONT-SIZE: smaller; FONT-FAMILY: Verdana, Arial, Helvetica, sans-serif } .sci_dirBtn { BACKGROUND-COLOR: #7aa544 } .sci_dirBtnHov { COLOR: #ffffff; BACKGROUND-COLOR: #1d6b2f } .blueBtn { BACKGROUND-COLOR: #0000aa } .blueBtnHov { COLOR: #0000aa; BACKGROUND-COLOR: #bad3f3 } .greenBtn { BACKGROUND-COLOR: #007703 } .greenBtnHov { COLOR: #007703; BACKGROUND-COLOR: #c2edc0 } .greyBtn { BACKGROUND-COLOR: #7f7f7f } .greyBtnHov { COLOR: #7f7f7f; BACKGROUND-COLOR: #e6e6e6 } .redBtn { BACKGROUND-COLOR: #990000 } .redBtnHov { COLOR: #990000; BACKGROUND-COLOR: #f7cece } .orangeBtn { BACKGROUND-COLOR: #cc5200 } .orangeBtnHov { COLOR: #cc5200; BACKGROUND-COLOR: #ffdfca } INPUT.btn { BORDER-RIGHT: #666666 1px solid; PADDING-RIGHT: 1px; BORDER-TOP: = #666666 1px solid; PADDING-LEFT: 1px; FONT-WEIGHT: bold; FONT-SIZE: = 12px; PADDING-BOTTOM: 0px; MARGIN: 0px 1px 0px 0px; VERTICAL-ALIGN: = middle; OVERFLOW: visible; BORDER-LEFT: #666666 1px solid; WIDTH: auto; = CURSOR: pointer; COLOR: #ffffff; PADDING-TOP: 0px; BORDER-BOTTOM: = #666666 1px solid; TEXT-DECORATION: none } INPUT.btnhov { BORDER-RIGHT: #666666 1px solid; PADDING-RIGHT: 1px; BORDER-TOP: = #666666 1px solid; PADDING-LEFT: 1px; FONT-WEIGHT: bold; FONT-SIZE: = 12px; PADDING-BOTTOM: 0px; MARGIN: 0px 1px 0px 0px; VERTICAL-ALIGN: = middle; OVERFLOW: visible; BORDER-LEFT: #666666 1px solid; WIDTH: auto; = CURSOR: pointer; PADDING-TOP: 0px; BORDER-BOTTOM: #666666 1px solid; = TEXT-DECORATION: none } A.articleBtn { BORDER-RIGHT: medium none; PADDING-RIGHT: 20px; BORDER-TOP: medium = none; PADDING-LEFT: 20px; FONT-WEIGHT: bold; FONT-SIZE: 12px; = PADDING-BOTTOM: 1px; MARGIN: 0px; BORDER-LEFT: medium none; CURSOR: = pointer; COLOR: #ffffff; PADDING-TOP: 1px; BORDER-BOTTOM: medium none; = BACKGROUND-COLOR: #999999; TEXT-DECORATION: none } A.articleBtn:hover { BORDER-RIGHT: medium none; PADDING-RIGHT: 20px; BORDER-TOP: medium = none; PADDING-LEFT: 20px; FONT-WEIGHT: bold; FONT-SIZE: 12px; = PADDING-BOTTOM: 1px; MARGIN: 0px; BORDER-LEFT: medium none; CURSOR: = pointer; COLOR: #ffffff; PADDING-TOP: 1px; BORDER-BOTTOM: medium none; = BACKGROUND-COLOR: #999999; TEXT-DECORATION: none } A.articleBtnActive { CURSOR: default; BACKGROUND-COLOR: #7aa544; TEXT-DECORATION: none } A.articleBtnActive:hover { CURSOR: default; BACKGROUND-COLOR: #7aa544; TEXT-DECORATION: none } A.articleBtn:hover { BACKGROUND-COLOR: #7aa544 } A.btn { BORDER-RIGHT: #666666 1px solid; PADDING-RIGHT: 5px; BORDER-TOP: = #666666 1px solid; PADDING-LEFT: 5px; FONT-WEIGHT: bold; FONT-SIZE: = 12px; PADDING-BOTTOM: 1px; MARGIN: 0px 1px 0px 0px; VERTICAL-ALIGN: = middle; BORDER-LEFT: #666666 1px solid; COLOR: #ffffff; PADDING-TOP: = 1px; BORDER-BOTTOM: #666666 1px solid; FONT-FAMILY: arial, verdana, = helvetica, sans-serif; HEIGHT: 1.17em; TEXT-DECORATION: none } A.btn:hover { BORDER-RIGHT: #666666 1px solid; PADDING-RIGHT: 5px; BORDER-TOP: = #666666 1px solid; PADDING-LEFT: 5px; FONT-WEIGHT: bold; FONT-SIZE: = 12px; PADDING-BOTTOM: 1px; MARGIN: 0px 1px 0px 0px; VERTICAL-ALIGN: = middle; BORDER-LEFT: #666666 1px solid; COLOR: #ffffff; PADDING-TOP: = 1px; BORDER-BOTTOM: #666666 1px solid; FONT-FAMILY: arial, verdana, = helvetica, sans-serif; HEIGHT: 1.17em; TEXT-DECORATION: none } A.sci_dirBtn { BACKGROUND-COLOR: #7aa544 } A.sci_dirBtn:hover { COLOR: #ffffff; BACKGROUND-COLOR: #1d6b2f } A.blueBtn { BACKGROUND-COLOR: #0000aa } A.blueBtn:hover { COLOR: #0000aa; BACKGROUND-COLOR: #bad3f3 } A.greenBtn { BACKGROUND-COLOR: #007703 } A.greenBtn:hover { COLOR: #007703; BACKGROUND-COLOR: #c2edc0 } A.greyBtn { BACKGROUND-COLOR: #7f7f7f } A.greyBtn:hover { COLOR: #7f7f7f; BACKGROUND-COLOR: #e6e6e6 } A.redBtn { BACKGROUND-COLOR: #990000 } A.redBtn:hover { COLOR: #990000; BACKGROUND-COLOR: #f7cece } A.orangeBtn { BACKGROUND-COLOR: #cc5200 } A.orangeBtn:hover { COLOR: #cc5200; BACKGROUND-COLOR: #ffdfca } A.noul:hover { TEXT-DECORATION: none } .tableFmBg { PADDING-RIGHT: 0px; BORDER-TOP: #8cc919 2px solid; PADDING-LEFT: 0px; = PADDING-BOTTOM: 0px; MARGIN: 0px; PADDING-TOP: 0px; BORDER-BOTTOM: = #8cc919 2px solid; BACKGROUND-COLOR: #efefef } .sci_dirBorder { BORDER-TOP: #8cc919 2px solid; BORDER-BOTTOM: #8cc919 2px solid } .blueBorder { BORDER-TOP: #0000aa 2px solid; BORDER-BOTTOM: #0000aa 2px solid } .greenBorder { BORDER-TOP: #007703 2px solid; BORDER-BOTTOM: #007703 2px solid } .greyBorder { BORDER-TOP: #7f7f7f 2px solid; BORDER-BOTTOM: #7f7f7f 2px solid } .redBorder { BORDER-TOP: #990000 2px solid; BORDER-BOTTOM: #990000 2px solid } .orangeBorder { BORDER-TOP: #cc5200 2px solid; BORDER-BOTTOM: #cc5200 2px solid } .sci_dirBorderBtm { BORDER-BOTTOM: #8cc919 2px solid } .blueBorderBtm { BORDER-BOTTOM: #0000aa 2px solid } .greenBorderBtm { BORDER-BOTTOM: #007703 2px solid } .greyBorderBtm { BORDER-BOTTOM: #7f7f7f 2px solid } .redBorderBtm { BORDER-BOTTOM: #990000 2px solid } .orangeBorderBtm { BORDER-BOTTOM: #cc5200 2px solid } .tableFmBgGray { FONT-SIZE: 1em; FONT-FAMILY: arial, helvetica, sans-serif; = BACKGROUND-COLOR: #efefef } .tableFmBgGrayNoCell { PADDING-RIGHT: 5px; PADDING-LEFT: 5px; FONT-SIZE: 1em; PADDING-BOTTOM: = 5px; MARGIN: 5px; PADDING-TOP: 5px; FONT-FAMILY: arial, helvetica, = sans-serif; BACKGROUND-COLOR: #efefef } .tableFmBgPad { MARGIN-TOP: 5px; MARGIN-BOTTOM: 5px; PADDING-BOTTOM: 5px; PADDING-TOP: = 5px; BACKGROUND-COLOR: #efefef } .tableFmCell { PADDING-LEFT: 10px; FONT-WEIGHT: bold; PADDING-BOTTOM: 3px; = MARGIN-LEFT: 10px; VERTICAL-ALIGN: top; PADDING-TOP: 3px; TEXT-ALIGN: = right } .tableFmCellTop { FONT-WEIGHT: bold; VERTICAL-ALIGN: top; PADDING-TOP: 0px; TEXT-ALIGN: = right } .tableFmCellAbove { PADDING-RIGHT: 5px; PADDING-LEFT: 5px; FONT-WEIGHT: bold; = PADDING-BOTTOM: 5px; PADDING-TOP: 5px; TEXT-ALIGN: left } .tableFmCellL { PADDING-LEFT: 10px; FONT-WEIGHT: bold; PADDING-BOTTOM: 3px; = MARGIN-LEFT: 10px; VERTICAL-ALIGN: top; PADDING-TOP: 3px; TEXT-ALIGN: = left } .tableFmDivider { PADDING-RIGHT: 0px; PADDING-LEFT: 0px; PADDING-BOTTOM: 0px; MARGIN: = 0px; WIDTH: 100%; PADDING-TOP: 0px; HEIGHT: 2px; BACKGROUND-COLOR: = #ffffff } .redtableFmDivider { PADDING-RIGHT: 0px; PADDING-LEFT: 0px; PADDING-BOTTOM: 0px; MARGIN: = 0px; WIDTH: 100%; PADDING-TOP: 0px; HEIGHT: 2px; BACKGROUND-COLOR: = #990000 } .SDtxtLarger { FONT-WEIGHT: bold; FONT-SIZE: larger; FONT-FAMILY: arial, helvetica, = sans-serif } .SDtxtSmall { FONT-SIZE: xx-small; COLOR: #000000; FONT-FAMILY: arial, helvetica, = sans-serif } .SDtxtSmallBold { FONT-WEIGHT: bold; FONT-SIZE: 0.9em } .SDtxtNote { FONT-SIZE: 1em; COLOR: #000000; FONT-FAMILY: arial, helvetica, = sans-serif } .SDtxtNoteSmall { FONT-SIZE: 10px; COLOR: #000000; FONT-FAMILY: arial, helvetica, = sans-serif } .SDtxtPgTitleBlack { FONT-WEIGHT: bold; FONT-SIZE: 150%; COLOR: #000000; FONT-FAMILY: arial, = helvetica, sans-serif } A.errMsg { COLOR: #ff0000; TEXT-DECORATION: none } A.errMsg:hover { TEXT-DECORATION: underline } .boldText { FONT-WEIGHT: bold } .sci_dirColor { COLOR: #7aa544 } .blueColor { COLOR: #0000aa } .greenColor { COLOR: #007703 } .greyColor { COLOR: #7f7f7f } .redColor { COLOR: #990000 } .orangeColor { COLOR: #cc5200 } .tablePlainWhite { PADDING-RIGHT: 0px; PADDING-LEFT: 0px; PADDING-BOTTOM: 0px; = PADDING-TOP: 0px; BACKGROUND-COLOR: #ffffff } .inputBox { BORDER-RIGHT: #666666 1px solid; BORDER-TOP: #666666 1px solid; = FONT-SIZE: 1em; VERTICAL-ALIGN: top; BORDER-LEFT: #666666 1px solid; = BORDER-BOTTOM: #666666 1px solid; BACKGROUND-COLOR: #ffffff } A.tabActive { BORDER-RIGHT: medium none; PADDING-RIGHT: 20px; BORDER-TOP: medium = none; PADDING-LEFT: 20px; FONT-WEIGHT: bold; FONT-SIZE: 1em; = PADDING-BOTTOM: 1px; MARGIN: 0px 2px 0px 0px; BORDER-LEFT: medium none; = COLOR: #ffffff; PADDING-TOP: 1px; BORDER-BOTTOM: medium none; = TEXT-DECORATION: none } A.tabActive:hover { BORDER-RIGHT: medium none; PADDING-RIGHT: 20px; BORDER-TOP: medium = none; PADDING-LEFT: 20px; FONT-WEIGHT: bold; FONT-SIZE: 1em; = PADDING-BOTTOM: 1px; MARGIN: 0px 2px 0px 0px; BORDER-LEFT: medium none; = COLOR: #ffffff; PADDING-TOP: 1px; BORDER-BOTTOM: medium none; = TEXT-DECORATION: none } A.tab { BORDER-RIGHT: medium none; PADDING-RIGHT: 20px; BORDER-TOP: medium = none; PADDING-LEFT: 20px; FONT-WEIGHT: bold; FONT-SIZE: 1em; = PADDING-BOTTOM: 1px; MARGIN: 0px 2px 0px 0px; BORDER-LEFT: medium none; = COLOR: #ffffff; PADDING-TOP: 1px; BORDER-BOTTOM: medium none; = BACKGROUND-COLOR: #999999; TEXT-DECORATION: none } A.tab:hover { BORDER-RIGHT: medium none; PADDING-RIGHT: 20px; BORDER-TOP: medium = none; PADDING-LEFT: 20px; FONT-WEIGHT: bold; FONT-SIZE: 1em; = PADDING-BOTTOM: 1px; MARGIN: 0px 2px 0px 0px; BORDER-LEFT: medium none; = COLOR: #ffffff; PADDING-TOP: 1px; BORDER-BOTTOM: medium none; = BACKGROUND-COLOR: #999999; TEXT-DECORATION: none } A.sci_dirTabActive { CURSOR: default; BACKGROUND-COLOR: #7aa544; TEXT-DECORATION: none } A.sci_dirTabActive:hover { CURSOR: default; BACKGROUND-COLOR: #7aa544; TEXT-DECORATION: none } A.sci_dirTab:hover { BACKGROUND-COLOR: #7aa544 } A.blueTabActive { CURSOR: default; BACKGROUND-COLOR: #0000aa; TEXT-DECORATION: none } A.blueTabActive:hover { CURSOR: default; BACKGROUND-COLOR: #0000aa; TEXT-DECORATION: none } A.blueTab:hover { BACKGROUND-COLOR: #0000aa } A.greenTabActive { CURSOR: default; BACKGROUND-COLOR: #007703; TEXT-DECORATION: none } A.greenTabActive:hover { CURSOR: default; BACKGROUND-COLOR: #007703; TEXT-DECORATION: none } A.greenTab:hover { BACKGROUND-COLOR: #007703 } A.greyTabActive { CURSOR: default; BACKGROUND-COLOR: #7f7f7f; TEXT-DECORATION: none } A.greyTabActive:hover { CURSOR: default; BACKGROUND-COLOR: #7f7f7f; TEXT-DECORATION: none } A.greyTab:hover { BACKGROUND-COLOR: #7f7f7f } A.redTabActive { CURSOR: default; BACKGROUND-COLOR: #990000; TEXT-DECORATION: none } A.redTabActive:hover { CURSOR: default; BACKGROUND-COLOR: #990000; TEXT-DECORATION: none } A.redTab:hover { BACKGROUND-COLOR: #990000 } A.orangeTabActive { CURSOR: default; BACKGROUND-COLOR: #cc5200; TEXT-DECORATION: none } A.orangeTabActive:hover { CURSOR: default; BACKGROUND-COLOR: #cc5200; TEXT-DECORATION: none } A.orangeTab:hover { BACKGROUND-COLOR: #cc5200 } .textBoxMultipleRow { FONT-SIZE: 1em; FONT-FAMILY: Arial, Helvetica, sans-serif; = BACKGROUND-COLOR: #ffffff } A.nextPrev { FONT-SIZE: 1em; MARGIN: 0px 10px; WIDTH: 1%; COLOR: #ffffff; = TEXT-DECORATION: none } A.nextPrev:hover { FONT-SIZE: 1em; MARGIN: 0px 10px; WIDTH: 1%; COLOR: #ffffff; = TEXT-DECORATION: underline } INPUT.nextPrev { BORDER-RIGHT: medium none; PADDING-RIGHT: 1px; BORDER-TOP: medium none; = PADDING-LEFT: 1px; FONT-SIZE: 1em; PADDING-BOTTOM: 0px; MARGIN: 0px 5px; = VERTICAL-ALIGN: middle; OVERFLOW: visible; BORDER-LEFT: medium none; = WIDTH: auto; COLOR: #0000ee; PADDING-TOP: 0px; BORDER-BOTTOM: medium = none; BACKGROUND-COLOR: transparent; TEXT-DECORATION: none } INPUT.nextPrevHov { BORDER-RIGHT: medium none; PADDING-RIGHT: 1px; BORDER-TOP: medium none; = PADDING-LEFT: 1px; FONT-SIZE: 1em; PADDING-BOTTOM: 0px; MARGIN: 0px 5px; = VERTICAL-ALIGN: middle; OVERFLOW: visible; BORDER-LEFT: medium none; = WIDTH: auto; COLOR: #0000ee; PADDING-TOP: 0px; BORDER-BOTTOM: medium = none; BACKGROUND-COLOR: transparent; TEXT-DECORATION: underline } .smCaps { FONT-VARIANT: small-caps } .nowrap { WHITE-SPACE: nowrap } SPAN.hit { FONT-WEIGHT: bold; COLOR: #ff0000 } SPAN.bf { FONT-WEIGHT: bold } SPAN.it { FONT-STYLE: italic } SPAN.monospace { FONT-FAMILY: courier } SPAN.sans-serif { FONT-FAMILY: sans-serif } SPAN.underline { TEXT-DECORATION: underline } SPAN.pit { FONT-SIZE: 0.9em; COLOR: #737373; FONT-FAMILY: arial, verdana, = helvetica, sans-serif } .charImg { BORDER-RIGHT: medium none; BORDER-TOP: medium none; VERTICAL-ALIGN: = baseline; BORDER-LEFT: medium none; BORDER-BOTTOM: medium none } SPAN.cross-out { TEXT-DECORATION: line-through } .refersTable { FONT-SIZE: 1em; LINE-HEIGHT: 16px; FONT-FAMILY: arial, helvetica, = sans-serif; BACKGROUND-COLOR: #efefef } .refersLeftColumn { PADDING-LEFT: 5px; FONT-WEIGHT: bold; VERTICAL-ALIGN: top; WIDTH: 55px; = TEXT-ALIGN: right } .refersDivider { PADDING-RIGHT: 0px; PADDING-LEFT: 0px; PADDING-BOTTOM: 0px; MARGIN: = 0px; WIDTH: 100%; PADDING-TOP: 0px; HEIGHT: 2px; BACKGROUND-COLOR: = #ffffff } .articleTitle { FONT-WEIGHT: bold; FONT-SIZE: 18px; FONT-FAMILY: Arial, Helvetica, = sans-serif } .h1 { FONT-WEIGHT: bold; FONT-SIZE: 2em; FONT-FAMILY: = arial,verdana,helvetica,sans-serif } .h2 { FONT-WEIGHT: bold; FONT-SIZE: 1.3em; FONT-FAMILY: = arial,verdana,helvetica,sans-serif } .h3 { FONT-WEIGHT: bold; FONT-SIZE: 1.2em; FONT-FAMILY: = arial,verdana,helvetica,sans-serif } .h4 { FONT-WEIGHT: bold; FONT-SIZE: 1.1em; FONT-FAMILY: = arial,verdana,helvetica,sans-serif } .h5 { FONT-WEIGHT: bold; FONT-SIZE: 1em; FONT-FAMILY: = arial,verdana,helvetica,sans-serif } .ImageScroller { OVERFLOW: auto; WIDTH: 435px } .ImageScrollerPurchHist { OVERFLOW: auto; WIDTH: 455px } .ImageScrollerPurchCurr { OVERFLOW: auto; 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