From: Subject: Cell -- Morrison and Spradling Date: Mon, 3 Mar 2008 09:33:38 +0100 MIME-Version: 1.0 Content-Type: multipart/related; type="text/html"; boundary="----=_NextPart_000_0052_01C87D11.A93889F0" X-MimeOLE: Produced By Microsoft MimeOLE V6.00.2900.3198 This is a multi-part message in MIME format. ------=_NextPart_000_0052_01C87D11.A93889F0 Content-Type: text/html; charset="utf-8" Content-Transfer-Encoding: quoted-printable Content-Location: http://www.cell.com/content/article/abstract?uid=PIIS0092867408001396 =EF=BB=BF Cell -- Morrison and = Spradling
=20 Cancer Cell Cell Cell Host & = Microbe=20 Cell Stem Cell Chemistry & Biology = Current Biology = Developmental Cell = Immunity Molecular Cell Neuron Structure
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Copyright =C2=A9 2008 Cell Press. All rights=20 reserved.
Cell, Vol 132, 598-611, 22 February 2008

Review

Stem = Cells and=20 Niches: Mechanisms That Promote Stem Cell Maintenance throughout = Life

Sean J. Morrison1, and Allan C. Spradling2,

1 Howard = Hughes Medical=20 Institute, Life Sciences Institute, and Center for Stem Cell Biology, = University=20 of Michigan, Ann Arbor, MI, 48109-2216, USA2 Howard = Hughes Medical=20 Institute and Department of Embryology, Carnegie Institution, Baltimore, = MD=20 21218, USA

Corresponding author
Sean J. Morrison
seanjm@med.umich.edu

Corresponding author
Allan C. Spradling
spradling@ciwemb.edu

Summary


Niches are local tissue=20 microenvironments that maintain and regulate stem cells. Long-predicted = from=20 mammalian studies, these structures have recently been characterized = within=20 several invertebrate tissues using methods that reliably identify = individual=20 stem cells and their functional requirements. Although similar = single-cell=20 resolution has usually not been achieved in mammalian tissues, = principles likely=20 to govern the behavior of niches in diverse organisms are emerging. = Considerable=20 progress has been made in elucidating how the microenvironment promotes = stem=20 cell maintenance. Mechanisms of stem cell maintenance are key to the = regulation=20 of homeostasis and likely contribute to aging and tumorigenesis when = altered=20 during adulthood.

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BACKGROUND-COLOR: #003399; important:=20 } #popup_footer { BACKGROUND-COLOR: #003399; important:=20 } ------=_NextPart_000_0052_01C87D11.A93889F0 Content-Type: application/octet-stream Content-Transfer-Encoding: quoted-printable Content-Location: http://www.cell.com/webfiles/javascript/script.js function dropdown(list) { var item=3Dlist.options[list.selectedIndex].value; if (item !=3D "nowhere")=20 { top.location=3Ditem; } } =09 function openhelp() { window.open('/popup/help', 'helpwindow', = 'width=3D580,height=3D410,left=3D0,top=3D0,screenX=3D0,screenY=3D0,resiza= ble=3Dno,scrollbars=3Dyes'); self.name=3D"main"; } function opensearchhelp() { = window.open('/popup/help/Finding_Information.htm','helpwindow','width=3D5= 80,height=3D410,left=3D0,top=3D0,screenX=3D0,screenY=3D0,resizable=3Dno,s= crollbars=3Dyes'); self.name=3D"main"; } function statecodeselect(thisform, thislist, list1, list2, = countrycodefield, countrydescfield) { var theform =3D eval("document." + thisform); var thelist =3D eval("theform." + thislist); eval("theform." + list1 + ".selectedIndex=3D0"); eval("theform." + list2 + ".selectedIndex=3D0"); eval("theform.elements[\"" + countrycodefield + = "\"].value=3Dthelist.options[thelist.selectedIndex].text"); eval("theform.elements[\"" + countrydescfield + = "\"].value=3Dthelist.options[thelist.selectedIndex].text"); } function clearstates ( thisform, strControlNameRoot, strIdentifiers ) { eval ( "document." + thisform + "." + strControlNameRoot + = ".value=3D''" ); var colIdentifiers =3D strIdentifiers.split ( ";" ); for ( var iCounter =3D 0; iCounter < colIdentifiers.length; iCounter++ = ) eval ( "document." + thisform + "." + strControlNameRoot + "_" + = colIdentifiers[iCounter] + ".selectedIndex=3D0" ); } //function finalstate ( thisform, strControlNameRoot, strSelectRef, = strIdentifiers ) function final_state ( thisform, strSelectRef, strIdentifiers ) { var strCountryID =3D eval ( "document." + thisform + "." + strSelectRef = + ".options[document." + thisform + "." + strSelectRef + = ".selectedIndex].value" ); if ( strIdentifiers.indexOf ( strCountryID ) =3D=3D -1 || strCountryID = =3D=3D "0") return; //var strStateID =3D eval ( "document." + thisform + "." + = strControlNameRoot + "_" + strCountryID + ".options[document." + = thisform + "." + strControlNameRoot + "_" + strCountryID + = ".selectedIndex].value" ); //eval ( "document." + thisform + "." + strControlNameRoot ).value =3D = strStateID; } function finalstate ( thisform, strControlNameRoot, strSelectRef, = strIdentifiers ) //function finalstate ( thisform, strSelectRef, strIdentifiers ) { var strCountryID =3D eval ( "document." + thisform + "." + strSelectRef = + ".options[document." + thisform + "." + strSelectRef + = ".selectedIndex].value" ); if ( strIdentifiers.indexOf ( strCountryID ) =3D=3D -1 || strCountryID = =3D=3D "0") return; var strStateID =3D eval ( "document." + thisform + "." + = strControlNameRoot + "_" + strCountryID + ".options[document." + = thisform + "." + strControlNameRoot + "_" + strCountryID + = ".selectedIndex].value" ); eval ( "document." + thisform + "." + strControlNameRoot ).value =3D = strStateID; } function setstate ( thisform, strControlNameRoot, strIdentifier, = strSelectRef, strIdentifiers, evt ) { // The following line ensures that the event type can be picked up = across all IE and NN var eventType =3D evt.type || arguments.callee.caller.name || = window.event.type; if ( eventType.indexOf ( "on" ) !=3D 0 ) eventType =3D "on" + eventType; if ( strIdentifier !=3D "" ) { if ( eval ( "document." + thisform + "." + strSelectRef + = ".options[document." + thisform + "." + strSelectRef + = ".selectedIndex].value" ) !=3D strIdentifier ) { // This part will be hit if the user selects a drop down that isn't = the one that they should // The country has not been set, or is set to something other than = the state box that was selected // Need to do two things. Set the country to the relevant value and = clear the other boxes if ( eventType =3D=3D "onchange" ) { // First, clear the other drop downs' values var colIdentifiers =3D strIdentifiers.split ( ";" ); for ( var iCounter =3D 0; iCounter < colIdentifiers.length; = iCounter++ ) { if ( colIdentifiers[iCounter] !=3D strIdentifier ) eval ( "document." + thisform + "." + strControlNameRoot + "_" + = colIdentifiers[iCounter] + ".selectedIndex=3D0" ); } // Clear the text box eval ( "document." + thisform + "." + strControlNameRoot + = ".value=3D''" ); // Now set the country eval ( "document." + thisform + "." + strSelectRef + ".selectedIndex = =3D " + searchArray ( eval ( "document." + thisform + "." + strSelectRef = + ".options" ), strIdentifier ) ); } return; } return; } else { if ( ( eval ( "document." + thisform + "." + strSelectRef + = ".selectedIndex" ) =3D=3D 0 ) || ( eval ( "document." + thisform + "." + strSelectRef + = ".options[document." + thisform + "." + strSelectRef + = ".selectedIndex].value" ) =3D=3D 1224 ) || ( eval ( "document." + thisform + "." + strSelectRef + = ".options[document." + thisform + "." + strSelectRef + = ".selectedIndex].value" ) =3D=3D 1037 ) ) { // This part will be entered when the user selects the text box and = that doesn't match what's in the country box // What needs to be done is the country box is aligned and the other = state input boxes to be reset if ( eventType =3D=3D "onchange" ) { // First, clear the drop downs' values var colIdentifiers =3D strIdentifiers.split ( ";" ); for ( var iCounter =3D 0; iCounter < colIdentifiers.length; = iCounter++ ) eval ( "document." + thisform + "." + strControlNameRoot + "_" + = colIdentifiers[iCounter] + ".selectedIndex=3D0" ); // Now set the country // Should we set it to a country? Or should it be blank? Think it = should be set to blank. eval ( "document." + thisform + "." + strSelectRef + ".selectedIndex = =3D 0" ); } return; } return; } } // This function searches an array and returns the index at which the = search string occurs function searchArray ( arrToSearch, strSearchValue ) { for ( var iCounter =3D 0; iCounter < arrToSearch.length; iCounter++ ) if ( arrToSearch[iCounter].value =3D=3D strSearchValue ) return iCounter; return -1; } function sethidden(value,field,value2) { if(!value2) { if(field.options) value2=3Dfield.options[field.selectedIndex].value; else value2=3Dfield.value; } if(value2!=3D"") { field.form.elements[value].value=3Dvalue2; } } function setEcommerce2Hidden() { sethidden('titleid',orderdetails.addresstitleid); sethidden('firstname',orderdetails.addressfirstname); sethidden('lastname',orderdetails.addresslastname); sethidden('address1',orderdetails.addressstreet); sethidden('address2',orderdetails.addressstreet2); sethidden('city',orderdetails.addresscity); sethidden('state',orderdetails.addressstate); sethidden('countryid',orderdetails.addresscountry); sethidden('zip',orderdetails.addresspostcode); sethidden('telephone',orderdetails.addressphone); } // Logic for YOAS checkboxs function yoasValidate(ckb) { var cnt =3D document.journals.journal.length - 3; var bk =3D document.journals.backfile.length >=3D 3; =09 if (bk) { //alert("records " + cnt); for (j =3D cnt; j < (cnt+3); j++)=20 { if (eval("document.journals.journal[" + (j) + "].checked") =3D=3D = true)=20 { document.journals.journal[j].checked =3D false; if (j - cnt =3D=3D ckb - 1)=20 { document.journals.journal[j].checked =3D true; } } } } } function noCheckbox() { for (j =3D 0; j < checkbox_form.uid.length; j++) { =20 if (checkbox_form.uid[j].checked) {=20 return true;=20 } =20 } msg =3D "Please select at least one checkbox"; alert(msg); return (false); } function oneCheckbox() { for (j =3D 1; j < document.rangedate.TOC.length; j++) { =20 if (eval("document.rangedate.TOC[" + (j) + "].checked") = =3D=3D true) {=20 rangedate.TOC[0].checked =3D false =09 } if (document.rangedate.TOC[0].checked =3D=3D true) {=20 rangedate.TOC[j].checked =3D false =09 } } return; } function oneCheckbox1() { for (j =3D 1; j < document.rangedate.TOC.length; j++) { =20 if (document.rangedate.TOC[0].checked =3D=3D true) {=20 rangedate.TOC[j].checked =3D false =09 } } return; } function encode(string){ =20 return string; } function linkChange(linkRef){=09 var volIssue =3D document.all.volIssue.value; linkRef.href=3DlinkRef.href+"&volIssue=3D"+volIssue; return true; } function refreshParent() { window.opener.location.href =3D window.opener.location.href; if (window.opener.progressWindow) =09 { window.opener.progressWindow.close() } window.close(); } // This function displays the ad results. // It must be defined above the script that calls show_ads.js // to guarantee that it is defined when show_ads.js makes the = call-back. function google_ad_request_done(google_ads) { // Proceed only if we have ads to display! if (google_ads.length < 1 ) return; // Display ads in a table document.write("

"); // Print "Ads By Google" -- include link to Google feedback page if = available if (google_info.feedback_url) { document.write("Ads by Google"); } else { document.write("Ads By Google"); } =20 document.write("
"); // For text ads, display each ad in turn. // In this example, each ad goes in a new row in the table. if (google_ads[0].type =3D=3D 'text') { for(i =3D 0; i < google_ads.length; ++i) { =09 document.write("
" + "" + =20 google_ads[i].line1 + "
" + google_ads[i].line2 + " " + google_ads[i].line3 + "
" +=20 "" +=20 google_ads[i].visible_url + "
");=20 } } // For an image ad, display the image; there will be only one . if (google_ads[0].type =3D=3D 'image') { document.write("" + "" + ""); } // Finish up anything that needs finishing up document.write ("

"); } function loadFormValues() { //alert(document.getElementById("a2").value); document.getElementById("search_author2").value = =3Ddocument.getElementById("a2").value ; document.getElementById("search_author1").value = =3Ddocument.getElementById("a1").value ; document.getElementById("search_words_title").value = =3Ddocument.getElementById("title-abstract").value ; document.getElementById("search_yearfrom").value = =3Ddocument.getElementById("fromyear").value ; document.getElementById("search_yearto").value = =3Ddocument.getElementById("toyear").value ; if(document.getElementById("orand").value=3D=3D'or') { document.getElementById("or").checked=3Dtrue; } if(document.getElementById("keywordloc").value=3D=3D'anywhere') { document.getElementById("inanywhere").checked=3Dtrue; } if(document.getElementById("keywordoptions").value=3D=3D'any') { document.getElementById("kw_any").checked=3Dtrue; } if(document.getElementById("keywordoptions").value=3D=3D'exact') { document.getElementById("kw_exact").checked=3Dtrue; } if(document.getElementById("published").value=3D=3D'0') { document.getElementById("pub_week").checked=3Dtrue; } if(document.getElementById("published").value=3D=3D'1') { document.getElementById("pub_month").checked=3Dtrue; } if(document.getElementById("published").value=3D=3D'2') { document.getElementById("pub_6months").checked=3Dtrue; } if(document.getElementById("published").value=3D=3D'4') { document.getElementById("pub_all").checked=3Dtrue; } =20 if(document.getElementById("typearticle").value=3D=3D'research') { document.getElementById("search_research").checked=3Dtrue; } if(document.getElementById("typearticle").value=3D=3D'review') { document.getElementById("search_review").checked=3Dtrue; } if(document.getElementById("sortby").value=3D=3D'rank') { document.getElementById("search_rank").checked=3Dtrue; } =20 // For Journal Check boxes var journal=3Ddocument.getElementById("journal-name").value;=09 var length_cell=3Djournal.length; var = immuni=3Ddocument.getElementById("journal-name").value.match("Immunity");= var = neuron=3Ddocument.getElementById("journal-name").value.match("Neuron"); var = mcell=3Ddocument.getElementById("journal-name").value.match("Molecular");= var = struc=3Ddocument.getElementById("journal-name").value.match("Structure");= var = curbio=3Ddocument.getElementById("journal-name").value.match("Current"); var = dcell=3Ddocument.getElementById("journal-name").value.match("Developmenta= l"); var = ccell=3Ddocument.getElementById("journal-name").value.match("Cancer"); var = chbiol=3Ddocument.getElementById("journal-name").value.match("Chemistry")= ; var = cmet=3Ddocument.getElementById("journal-name").value.match("Metabolism");= var = chom=3Ddocument.getElementById("journal-name").value.match("Host"); var = stem=3Ddocument.getElementById("journal-name").value.match("Stem"); var cell =3D journal.substring(0,5); if(length_cell=3D=3D'4' || cell=3D=3D'Cell,'){ =09 document.getElementById("search_cell").checked =3D true; rangedate.TOC[0].checked =3D false; } if(immuni=3D=3D'Immunity'){ document.getElementById("search_immuni").checked =3D true; rangedate.TOC[0].checked =3D false; } if(neuron=3D=3D'Neuron'){ document.getElementById("search_neuron").checked =3D true; rangedate.TOC[0].checked =3D false; } =20 if(mcell=3D=3D'Molecular'){ document.getElementById("search_molcel").checked =3D true; rangedate.TOC[0].checked =3D false; } =20 if(struc=3D=3D'Structure'){ document.getElementById("search_foldes").checked =3D true; rangedate.TOC[0].checked =3D false; } =09 if(curbio=3D=3D'Current'){ document.getElementById("search_curbio").checked =3D true; rangedate.TOC[0].checked =3D false; } =09 if(dcell=3D=3D'Developmental'){ document.getElementById("search_dcell").checked =3D true; rangedate.TOC[0].checked =3D false; } =09 if(ccell=3D=3D'Cancer'){ document.getElementById("search_ccell").checked =3D true; rangedate.TOC[0].checked =3D false; } =09 if(chbiol=3D=3D'Chemistry'){ document.getElementById("search_chbiol").checked =3D true; rangedate.TOC[0].checked =3D false; } =09 if(cmet=3D=3D'Metabolism'){ document.getElementById("search_cmet").checked =3D true; rangedate.TOC[0].checked =3D false; } =20 if(chom=3D=3D'Host'){ document.getElementById("search_chom").checked =3D true; rangedate.TOC[0].checked =3D false; } =09 if(stem=3D=3D'Stem'){ document.getElementById("search_stem").checked =3D true; rangedate.TOC[0].checked =3D false; } =09 } ------=_NextPart_000_0052_01C87D11.A93889F0 Content-Type: application/octet-stream Content-Transfer-Encoding: quoted-printable Content-Location: https://ad.uk.doubleclick.net/adj/cell.rbi/content/article/abstract;abr=!webtv;sz=728x90;ord=19985? document.write('3D"Click');=0A= ------=_NextPart_000_0052_01C87D11.A93889F0 Content-Type: application/octet-stream Content-Transfer-Encoding: quoted-printable Content-Location: https://ad.uk.doubleclick.net/adj/cell.rbi/content/article/abstract;abr=!webtv;sz=336x280;ord=18276? document.write('\n